Dr Ulmar Grafe


Professor, Faculty of Science


Dr Ulmar Grafe is a Professor of Biology at Universiti Brunei Darussalam. He received his PhD from Cornell University (USA) and his Habilitation from Universität Würzburg (Germany). He has worked in Brunei at the Faculty of Science and the Institute of Biodiversity and Environmental Research since 2005. Prof Grafe is a tropical ecologist, behavioural ecologist, and animal physiologist with particular interests in understanding the functional role of amphibians in forest ecosystems, the sensory ecology of frog-biting midges, animal-plant mutualisms, and multimodal communication.


Functional diversity of amphibians
Sensory ecology of frog-biting midges
Community ecology of Bornean frogs
Animal-plant mutualisms


Interactions between bats and pitcher plants

Applications Invited

Physiological mechanisms mediating ultrasonic detection in amphibians

Applications Invited

Google Scholar Citations


Google Scholar h-index


Google Scholar i10-index


Meuche, I., A. Keller, H.H. Ahmadsah, N. Ahmad & T.U. Grafe 2016. Silent listeners: Can acoustic preferences of blood-sucking eavesdroppers (Diptera: Corethrellidae) predict the parasitism risk of their anuran hosts? Behavioral Ecology Early Access
Sickel W., T.U. Grafe, I. Meuche, I. Steffan-Dewenter, A. Keller 2016. Diet determines bacterial diversity and community structure in Bornean pitcher plants. Microbial Ecology First Online.
Bauer, U., K. Rembold & T. U. Grafe 2016. Carnivorous Nepenthes pitcher plants are a rich food source for a diverse vertebrate community. Journal of Natural History 50: 483-495.
Konopik, O., I. StCurrent Biology Covereffan-Dewenter, T.U. Grafe 2015. Effects of logging and oil palm expansion on stream frog communities on Borneo, Southeast Asia. Biotropica 47:636-643. Editor’s Choice 47(5).
Schöner, M.G, C.R. Schöner, R. Simon, T.U. Grafe, S. Puechmaille, LJ. Liaw, G. Kerth 2015. Bats are acoustically attracted to mutualistic carnivorous plants. Current Biology 25:1911-1916. (Highlighted as cover of journal)


Grant type: URG, Grant Number: 105, Project Title: Spatial ecology and movement patterns of amphibians and reptiles in rainforests of Brunei Darussalam, Investigators (PI/Co-PI): Dr Ulmar Grafe, Funding Details: yes, Start Date: 28/03/2010, End Date: 30/09/2011
Grant Type: URG, Grant Number: 193, Project Title: Interactions between carnivorous pitcher plants and woolly bats in Borneo, Investigators (PI/Co-PI): Dr Ulmar Grafe, Funding Details: yes, Start Date: 27/06/2011, End Date: 30/07/2016
Grant Type: BRC, Grant Number: 13, Project Title: Diversity and distribution of parasitic flies in Brunei Darussalam with implications for human health , Investigators (PI/Co-PI): Dr Ulmar Grafe, Dr Norhayati Ahmad, Funding Details: yes, Start Date: 25/10/2010, End Date: 25/10/2015
Grant Type: BRC, Grant Number: 11, Project Title: Studies of the diversity, distribution, bioacoustics and ecomorphology of Bats (Chiroptera), a major, highly diverse but understudied group of mammals in Brunei Darussalam, Investigators (PI/Co-PI): Dr Ulmar Grafe
Dr Hussein Taha (pending), Funding Details: yes, Start Date: 04/06/2015, End Date: 25/10/2015